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What does quilled mean?

Looking for the meaning or definition of the word quilled? Here's what it means.

Adjective
  1. having quills or similar structures
  2. (of a flower) having long, narrow petals or florets
  3. created through the process of quilling
  4. (of fabric) having small, rounded folds
  5. decorated with quillwork
Verb
  1. simple past tense and past participle of quill
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